W.A. Bentley

The hare, the tortoise and the crocodile: The ecology of angiosperm dominance, conifer persistence and fern filtering
Coomes, David A., R.B. Allen, W.A. Bentley, L.E. Burrows, Charles D. Canham, L. Fagan, David M. Forsyth, et al. 2005. “The Hare, The Tortoise And The Crocodile: The Ecology Of Angiosperm Dominance, Conifer Persistence And Fern Filtering”. J. Ecol. 93: 918-935. http://www.caryinstitute.org/reprints/Coomes_et_al_2005_JEcol-Hare_tortoise_and_crocodile.pdf.